[1
]Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge,
UK
[2
]Analytic and Translational Genetics Unit, Massachusetts General Hospital, Harvard
Medical School, Boston, Massachusetts, USA
[3
]Broad Institute of MIT and Harvard, Cambridge, Massachusetts, USA
[4
]Department of Gastroenterology and Hepatology, University of Groningen and University
Medical Center Groningen, Groningen, The Netherlands
[5
]Division of Gastroenterology, Hepatology and Nutrition, Department of Medicine, University
of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, USA
[6
]Department of Human Genetics, University of Pittsburgh Graduate School of Public Health,
Pittsburgh, Pennsylvania, USA
[7
]Cedars-Sinai F. Widjaja Inflammatory Bowel and Immunobiology Research Institute, Los
Angeles, California, USA
[8
]Medical Genetics Institute, Cedars-Sinai Medical Center, Los Angeles, California,
USA
[9
]Department of Genetics, Yale School of Medicine, New Haven, Connecticut, USA
[10
]Inflammatory Bowel Disease Research Group, Addenbrooke’s Hospital, University of Cambridge,
Cambridge, UK
[11
]Department of Health Studies, University of Chicago, Chicago, Illinois, USA
[12
]Department of Internal Medicine, Section of Digestive Diseases, Yale School of Medicine,
New Haven, Connecticut, USA
[13
]Center for Human Genetic Research, Massachusetts General Hospital, Harvard Medical
School, Boston, Massachusetts, USA
[14
]University of Maribor, Faculty of Medicine, Center for Human Molecular Genetics and
Pharmacogenomics, Maribor, Slovenia
[15
]University Medical Center Groningen, Department of Genetics, Groningen, The Netherlands
[16
]Department of Pathophysiology, Gastroenterology section, KU Leuven, Leuven, Belgium
[17
]Unit of Animal Genomics, Groupe Interdisciplinaire de Genoproteomique Appliquee (GIGA-R)
and Faculty of Veterinary Medicine, University of Liege, Liege, Belgium
[18
]Division of Gastroenterology, Centre Hospitalier Universitaire, Universite de Liege,
Liege, Belgium
[19
]Department of Medical and Molecular Genetics, King’s College London School of Medicine,
Guy’s Hospital, London, UK
[20
]Division of Rheumatology Immunology and Allergy, Brigham and Women’s Hospital, Boston,
Massachusetts, USA
[21
]Program in Medical and Population Genetics, Broad Institute, Cambridge, Massachusetts,
USA
[22
]Division of Genetics, Brigham and Women’s Hospital, Boston, Massachusetts, USA
[23
]Université de Montréal and the Montreal Heart Institute, Research Center, Montréal,
Québec, Canada
[24
]Department of Computer Science, New Jersey Institute of Technology, Newark, NJ 07102,
USA
[25
]Department of Gastroenterology & Hepatology, Digestive Disease Institute, Cleveland
Clinic, Cleveland, Ohio
[26
]Department of Pathobiology, Lerner Research Institute, Cleveland Clinic, Cleveland,
Ohio, USA
[27
]Peninsula College of Medicine and Dentistry, Exeter, UK
[28
]Erasmus Hospital, Free University of Brussels, Department of Gastroenterology, Brussels,
Belgium
[29
]Massachusetts General Hospital, Harvard Medical School, Gastroenterology Unit, Boston,
Massachusetts, USA
[30
]Viborg Regional Hospital, Medical Department, Viborg, Denmark
[31
]Inflammatory Bowel Disease Service, Department of Gastroenterology and Hepatology,
Royal Adelaide Hospital, and School of Medicine, University of Adelaide, Adelaide,
Australia
[32
]Institute of Clinical Molecular Biology, Christian-Albrechts-University, Kiel, Germany
[33
]Department of Gastroenterology and Hepatology, Flinders Medical Centre and School
of Medicine, Flinders University, Adelaide, Australia
[34
]Division of Gastroenterology, McGill University Health Centre, Royal Victoria Hospital,
Montréal, Québec, Canada
[35
]Department of Medicine II, University Hospital Munich-Grosshadern, Ludwig-Maximilians-University,
Munich, Germany
[36
]Department of Gastroenterology, Charit, Campus Mitte, UniversitŠtsmedizin Berlin,
Berlin, Germany
[37
]Department of Genetics and Genomic Sciences, Mount Sinai School of Medicine, New York
City, New York, USA
[38
]Department of Genomics, Life & Brain Center, University Hospital Bonn, Bonn, Germany
[39
]Department of Biosciences and Nutrition, Karolinska Institutet, Stockholm, Sweden
[40
]Department of Pediatrics, Cedars Sinai Medical Center, Los Angeles, California, USA
[41
]Torbay Hospital, Department of Gastroenterology, Torbay, Devon, UK
[42
]School of Medical Sciences, Faculty of Medical & Health Sciences, The University of
Auckland, Auckland, New Zealand
[43
]University of Groningen, University Medical Center Groningen, Department of Genetics,
Groningen, The Netherlands
[44
]Department of Medicine, University of Otago, Christchurch, New Zealand
[45
]Department of Gastroenterology, Christchurch Hospital, Christchurch, New Zealand
[46
]Institute of Genetic Epidemiology, Helmholtz Zentrum München - German Research Center
for Environmental Health, Neuherberg, Germany
[47
]St Mark’s Hospital, Watford Road, Harrow, Middlesex, HA1 3UJ
[48
]Nottingham Digestive Diseases Centre, Queens Medical Centre, Nottingham NG7 1AW, UK
[49
]Research Institute of Internal Medicine, Oslo University Hospital Rikshospitalet,
Oslo, Norway
[50
]Kaunas University of Medicine, Department of Gastroenterology, Kaunas, Lithuania
[51
]Department of Pediatrics, Emory University School of Medicine, Atlanta, Georgia, USA
[52
]Unit of Gastroenterology, Istituto di Ricovero e Cura a Carattere Scientifico-Casa
Sollievo della Sofferenza (IRCCS-CSS) Hospital, San Giovanni Rotondo, Italy
[53
]Ghent University Hospital, Department of Gastroenterology and Hepatology, Ghent, Belgium
[54
]School of Medicine and Pharmacology, The University of Western Australia, Fremantle,
Australia
[55
]Gastrointestinal Unit, Molecular Medicine Centre, University of Edinburgh, Western
General Hospital, Edinburgh, UK
[56
]Department of Gastroenterology, The Townsville Hospital, Townsville, Australia
[57
]Institute of Human Genetics, Newcastle University, Newcastle upon Tyne, UK
[58
]Department of Medicine, Ninewells Hospital and Medical School, Dundee, UK
[59
]Genetic Medicine, MAHSC, University of Manchester, Manchester, UK
[60
]Academic Medical Center, Department of Gastroenterology, Amsterdam, The Netherlands
[61
]University of Maribor, Faculty for Chemistry and Chemical Engineering, Maribor, Slovenia
[62
]King’s College London School of Medicine, Guy’s Hospital, Department of Medical and
Molecular Genetics, London, UK
[63
]Royal Hospital for Sick Children, Paediatric Gastroenterology and Nutrition, Glasgow,
UK
[64
]Guy’s & St. Thomas’ NHS Foundation Trust, St. Thomas’ Hospital, Department of Gastroenterology,
London, UK
[65
]Department of Gastroenterology, Hospital Cl’nic/Institut d’Investigaci— Biomdica August
Pi i Sunyer (IDIBAPS), Barcelona, Spain
[66
]Centro de Investigaci—n Biomdica en Red de Enfermedades Hep‡ticas y Digestivas (CIBER
EHD), Barcelona, Spain
[67
]Christian-Albrechts-University, Institute of Clinical Molecular Biology, Kiel, Germany
[68
]Department for General Internal Medicine, Christian-Albrechts-University, Kiel, Germany
[69
]Inflammatory Bowel Diseases, Genetics and Computational Biology, Queensland Institute
of Medical Research, Brisbane, Australia
[70
]Norfolk and Norwich University Hospital
[71
]Department of Gastroenterology, Leiden University Medical Center, Leiden, The Netherlands
[72
]Child Life and Health, University of Edinburgh, Edinburgh, Scotland, UK
[73
]Institute of Human Genetics and Department of Neurology, Technische Universität München,
Munich, Germany
[74
]Center for Computational and Integrative Biology, Massachusetts General Hospital,
Boston, Massachusetts, USA
[75
]Department for General Internal Medicine, Christian-Albrechts-University, Kiel, Germany
[76
]Department of Biostatistics, School of Public Health, Yale University, New Haven,
Connecticut, USA
[78
]Mount Sinai Hospital Inflammatory Bowel Disease Centre, University of Toronto, Toronto,
Ontario, Canada
[79
]Azienda Ospedaliero Universitaria (AOU) Careggi, Unit of Gastroenterology SOD2, Florence,
Italy
[80
]Center for Applied Genomics, The Children’s Hospital of Philadelphia, Philadelphia,
Pennsylvania, USA
[81
]Department of Pediatrics, Center for Pediatric Inflammatory Bowel Disease, The Children’s
Hospital of Philadelphia, Philadelphia, Pennsylvania, USA
[82
]Meyerhoff Inflammatory Bowel Disease Center, Department of Medicine, School of Medicine,
and Department of Epidemiology, Bloomberg School of Public Health, Johns Hopkins University,
Baltimore, Maryland, USA
[83
]Department of Gastroenterology, Royal Brisbane and Womens Hospital, and School of
Medicine, University of Queensland, Brisbane, Australia
[84
]Inflammatory Bowel Disease Research Group, Addenbrooke’s Hospital, University of Cambridge,
Cambridge, UK
[85
]Division of Gastroenterology, University Hospital Gasthuisberg, Leuven, Belgium