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      An Evolutionary Model of Cooperation, Fairness and Altruistic Punishment in Public Good Games

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      PLoS ONE
      Public Library of Science

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          Abstract

          We identify and explain the mechanisms that account for the emergence of fairness preferences and altruistic punishment in voluntary contribution mechanisms by combining an evolutionary perspective together with an expected utility model. We aim at filling a gap between the literature on the theory of evolution applied to cooperation and punishment, and the empirical findings from experimental economics. The approach is motivated by previous findings on other-regarding behavior, the co-evolution of culture, genes and social norms, as well as bounded rationality. Our first result reveals the emergence of two distinct evolutionary regimes that force agents to converge either to a defection state or to a state of coordination, depending on the predominant set of self- or other-regarding preferences. Our second result indicates that subjects in laboratory experiments of public goods games with punishment coordinate and punish defectors as a result of an aversion against disadvantageous inequitable outcomes. Our third finding identifies disadvantageous inequity aversion as evolutionary dominant and stable in a heterogeneous population of agents endowed initially only with purely self-regarding preferences. We validate our model using previously obtained results from three independently conducted experiments of public goods games with punishment.

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          The evolution of cooperation.

          Cooperation in organisms, whether bacteria or primates, has been a difficulty for evolutionary theory since Darwin. On the assumption that interactions between pairs of individuals occur on a probabilistic basis, a model is developed based on the concept of an evolutionarily stable strategy in the context of the Prisoner's Dilemma game. Deductions from the model, and the results of a computer tournament show how cooperation based on reciprocity can get started in an asocial world, can thrive while interacting with a wide range of other strategies, and can resist invasion once fully established. Potential applications include specific aspects of territoriality, mating, and disease.
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            Via freedom to coercion: the emergence of costly punishment.

            In human societies, cooperative behavior in joint enterprises is often enforced through institutions that impose sanctions on defectors. Many experiments on so-called public goods games have shown that in the absence of such institutions, individuals are willing to punish defectors, even at a cost to themselves. Theoretical models confirm that social norms prescribing the punishment of uncooperative behavior are stable-once established, they prevent dissident minorities from spreading. But how can such costly punishing behavior gain a foothold in the population? A surprisingly simple model shows that if individuals have the option to stand aside and abstain from the joint endeavor, this paves the way for the emergence and establishment of cooperative behavior based on the punishment of defectors. Paradoxically, the freedom to withdraw from the common enterprise leads to enforcement of social norms. Joint enterprises that are compulsory rather than voluntary are less likely to lead to cooperation.
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              The long-run benefits of punishment.

              Experiments have shown that punishment enhances socially beneficial cooperation but that the costs of punishment outweigh the gains from cooperation. This challenges evolutionary models of altruistic cooperation and punishment, which predict that punishment will be beneficial. We compared 10- and 50-period cooperation experiments. With the longer time horizon, punishment is unambiguously beneficial.
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                Author and article information

                Contributors
                Role: Editor
                Journal
                PLoS One
                PLoS ONE
                plos
                plosone
                PLoS ONE
                Public Library of Science (San Francisco, USA )
                1932-6203
                2013
                19 November 2013
                : 8
                : 11
                : e77041
                Affiliations
                [1 ]Chair of Entrepreneurial Risks, Department of Management, Technology and Economics, ETH-Zurich, Zurich, Switzerland
                [2 ]Swiss Finance Institute, c/o University of Geneva, Geneva, Switzerland
                University of Zaragoza, Spain
                Author notes

                Competing Interests: The authors have declared that no competing interests exist.

                Analyzed the data: MH DS. Wrote the paper: MH DS. Formulated the problem: MH DS. Developed the theory: MH DS. Analyzed the experimental data: MH DS.

                Article
                PONE-D-13-19607
                10.1371/journal.pone.0077041
                3834069
                24260101
                ae41dfeb-4925-4e60-8fa7-54f45b4e7731
                Copyright @ 2013

                This is an open-access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.

                History
                : 12 May 2013
                : 5 September 2013
                Page count
                Pages: 13
                Funding
                The authors have no support or funding to report.
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                Research Article

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