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      Kikuyu-based pasture for dairy production: a review

      , , ,
      Crop and Pasture Science
      CSIRO Publishing

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          Abstract

          The amount of pasture grown and converted to animal product is closely linked with the profitability of pasture-based systems. Kikuyu (Pennisetum clandestinum Hochst. ex Chiov.) is the predominant C4 grass in coastal Australian beef and dairy systems. These kikuyu-based production systems face several key challenges to achieving high levels of productivity. In this review, we bring together the literature to highlight the opportunities for closing the gap between current and potential utilisation and for increasing dairy production from kikuyu-based pastures. More specifically, we highlight the significant gains that can be made on kikuyu-based commercial farms based on a conceptual model to show where the main losses originate, namely input and grazing management. The physical limitations associated with kikuyu for dairy systems are also presented, such as the relatively higher content of cell wall and lower content of water-soluble carbohydrates, together with nutrient imbalances relative to other grass species. Together, these limitations clearly indicate the need of supplying cows with supplements (particularly grain-based concentrates) to achieve moderate to high milk yield per cow. To achieve this without compromising pasture utilisation, dairy producers farming on kikuyu-based pastures need to use relatively greater stocking rates to generate enough demand of feed that can be used to align rate of pasture intake with rate of pasture growth, creating enough deficit of feed per cow to justify the addition of supplementary feed without impinging on pasture utilisation. The variability that exists between cows in kikuyu dry matter and neutral detergent fibre intake is also highlighted in this review, opening up new avenues of research that may allow significant productivity gains for kikuyu-based dairy farming in the future.

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          Predicting intake and digestibility using mathematical models of ruminal function.

          D Mertens (1987)
          Intake and digestibility of feeds by ruminants are influenced by characteristics of the feed, animal and feeding situation. Integration of these characteristics in mathematical models is critical to future progress in forage evaluation and optimal formulation of diets for ruminants. The physiological and physical theories of intake regulation can be described by simple mathematical equations. These equations indicate that intake is a linear function of animal characteristics, such as body weight and production level, and a reciprocal function of feed characteristics, such as fill effect and energy content. Theoretical equations were developed to predict intake when the neutral detergent fiber and energy content of the diet and the energy requirements of the animal are known. The theoretical model also can be used to predict the maximum intake that will maintain a given level of animal production by solving the physiological and physical intake equations at their intersection. Psychogenic intake regulation, which is related to the animal's behavioral response to factors not related to physiological or physical characteristics, can be described mathematically as a multiplier. Digestibility can be predicted by summing the contents of ideal nutritive entities in feeds, which have true digestibilities near 100%, subtracting their associated endogenous losses and adding the variable digestible fiber content. Steady-state models indicate fractional rates of digestion and passage can be used to define ideal nutritive entities and predict digestibility over a range of kinetic characteristics. The steady-state solutions are particularly useful in understanding and predicting the depression in digestibility associated with changes in rates of passage at high levels of feed intake.
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            Invited review: production and digestion of supplemented dairy cows on pasture.

            Literature with data from dairy cows on pasture was reviewed to evaluate the effects of supplementation on intake, milk production and composition, and ruminal and postruminal digestion. Low dry matter intake (DMI) of pasture has been identified as a major factor limiting milk production by high producing dairy cows. Pasture DMI in grazing cows is a function of grazing time, biting rate, and bite mass. Concentrate supplementation did not affect biting rate (58 bites/min) or bite mass (0.47 g of DM/bite) but reduced grazing time 12 min/d per kilogram of concentrate compared with unsupplemented cows (574 min/d). Substitution rate, or the reduction in pasture DMI per kilogram of concentrate, is a factor which may explain the variation in milk response to supplementation. A negative relationship exists between substitution rate and milk response; the lower the substitution rate the higher the milk response to supplements. Milk production increases linearly as the amount of concentrate increases from 1.2 to 10 kg DM/d, with an overall milk response of 1 kg milk/kg concentrate. Compared with pasture-only diets, increasing the amount of concentrate supplementation up to 10 kg DM/d increased total DMI 24%, milk production 22%, and milk protein percentage 4%, but reduced milk fat percentage 6%. Compared with dry ground corn, supplementation with nonforage fiber sources or processed corn did not affect total DMI, milk production, or milk composition. Replacing ruminal degradable protein sources with ruminal undegradable protein sources in concentrates did not consistently affect milk production or composition. Forage supplementation did not affect production when substitution rate was high. Fat supplementation increased milk production by 6%, without affecting milk fat and protein content. Increasing concentrate from 1.1 to 10 kg DM/d reduced ruminal pH 0.08 and NH3-N concentration 6.59 mg/dl, compared with pasture-only diets. Replacing dry corn by high moisture corn, steam-flaked or steam-rolled corn, barley, or fiber-based concentrates reduced ruminal NH3-N concentration 4.36 mg/dl. Supplementation did not affect in situ pasture digestion, except for a reduction in rate of degradation when high amounts of concentrate were supplemented. Supplementation with energy concentrates reduced digestibility of neutral detergent fiber and intake of N but did not affect digestibility of organic matter or flow of microbial N.
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              Performance and nutrient intake of high producing Holstein cows consuming pasture or a total mixed ration.

              We compared the intakes of nutrients by high producing Holstein cows consuming pasture or a full nutrient positive control ration (total mixed ration; TMR) and identified nutrients that limited the milk production of cows consuming the high quality pasture. Cows (n = 8) were adapted to an all pasture diet by incrementally reducing the amount of TMR fed over a 4-wk period. A control group of cows (n = 8) remained in confinement and was fed a TMR. The performance of grazing cows differed significantly from that of cows fed the TMR in dry matter (DM) intake (19.0 vs. 23.4 kg/d of DM), milk production (29.6 vs. 44.1 kg/d), milk protein content (2.61% vs. 2.80%), live weight (562 vs. 597 kg), and body condition score (2.0 vs. 2.5). The high quality of the pasture permitted cows to consume the same daily intakes of neutral detergent fiber and crude protein (kilograms per day) as cows fed the TMR, but the pasture provided 19% less DM, organic matter, and net energy for lactation. Predictions using National Research Council estimates and the Cornell Net Carbohydrate and Protein System model indicated that the supply of metabolizable energy was first-limiting for the milk production of cows consuming high quality pasture rather than the supply of metabolizable protein or amino acids. Although a daily intake of 19 kg of DM was achieved on spring pasture, the significant mobilization of energy reserves indicated that supplemental energy is required to achieve milk production greater than 30 kg/d from high producing Holstein cows on intensive grazing systems.
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                Author and article information

                Journal
                Crop and Pasture Science
                Crop Pasture Sci.
                CSIRO Publishing
                1836-0947
                2014
                2014
                : 65
                : 8
                : 787
                Article
                10.1071/CP13414
                3e377da7-d42b-45ed-a962-aab619200347
                © 2014
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